| Main Ref. | Conand, C., 1998 | 
|---|---|
| Remarks | A reef species mostly found in shallow areas from near the surface to a depth of 15 m (Ref. 122). Found on sandy areas (Ref. 115900) and generally occurs on reef flats and upper slopes. Populations reaching high densities on hard substrates, with a mean of around 0.04 per square meter. Shallow water populations are generally smaller in size than those found in deeper waters. Biology poorly known (Ref. 122). | 
| Marine - Neritic | Marine - Oceanic | Brackishwater | Freshwater | |
|---|---|---|---|---|
| Marine zones / Brackish and freshwater bodies | 
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| Substrate | Benthic: mobile; Soft Bottom: sand; Hard Bottom; | 
|---|---|
| Substrate Ref. | |
| Special habitats | Coral Reefs: reef slope/drop-offs; reef flats; | 
| Special habitats Ref. | 
| Ref. | Conand, C., 1998 | 
|---|---|
| Associations | symbiosis; | 
| Associated with | Carapus mourlani (fish) (Ref. 102254); Periclimenes imperator (shrimp) (Ref. 102275). | 
| Association remarks | |
| Parasitism | 
| Feeding type | mainly plants/detritus (troph. 2-2.19) | 
|---|---|
| Feeding type Ref. | |
| Feeding habit | browsing on substrate | 
| Feeding habit Ref. | 
| Estimation method | original sample | unfished population | Remark | ||
|---|---|---|---|---|---|
| Troph | s.e. | Troph | s.e. | ||
| From diet composition | |||||
| From individual food items | |||||
| Ref. | |||||