| Main Ref. | Roper, C.F.E., M.J. Sweeney and C.E. Nauen, 1984 | 
|---|---|
| Remarks | Sometimes found burrowing in the substrate (Ref. 275). | 
| Marine - Neritic | Marine - Oceanic | Brackishwater | Freshwater | |
|---|---|---|---|---|
| Marine zones / Brackish and freshwater bodies | 
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 | 
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| Substrate | Benthic: mobile; demersal; Soft Bottom: sand; | 
|---|---|
| Substrate Ref. | Roper, C.F.E., M.J. Sweeney and C.E. Nauen, 1984 | 
| Special habitats | Other habitats: burrows; | 
| Special habitats Ref. | Carpenter, K.E. and V.H. Niem, 1998 | 
| Ref. | Roper, C.F.E., M.J. Sweeney and C.E. Nauen, 1984 | 
|---|---|
| Associations | |
| Associated with | |
| Association remarks | |
| Parasitism | 
| Feeding type | mainly animals (troph. 2.8 and up) | 
|---|---|
| Feeding type Ref. | |
| Feeding habit | hunting macrofauna (predator) | 
| Feeding habit Ref. | 
| Estimation method | original sample | unfished population | Remark | ||
|---|---|---|---|---|---|
| Troph | s.e. | Troph | s.e. | ||
| From diet composition | |||||
| From individual food items | 3.39 | 0.50 | Trophic level estimated from a number of food items using a randomized resampling routine. | ||
| Ref. | |||||