| Main Ref. | Austin, W.C., 1999 | 
|---|---|
| Remarks | Mainly lives in the littoral zone but found to a depth of 90 m (Ref.123216). | 
| Marine - Neritic | Marine - Oceanic | Brackishwater | Freshwater | |
|---|---|---|---|---|
| Marine zones / Brackish and freshwater bodies | 
 | 
 | 
 | 
 | 
| Substrate | Soft Bottom: sand; | 
|---|---|
| Substrate Ref. | Miyake, S., 1957 | 
| Special habitats | |
| Special habitats Ref. | 
| Ref. | |
|---|---|
| Associations | |
| Associated with | |
| Association remarks | |
| Parasitism | 
| Feeding type | |
|---|---|
| Feeding type Ref. | |
| Feeding habit | |
| Feeding habit Ref. | 
| Estimation method | original sample | unfished population | Remark | ||
|---|---|---|---|---|---|
| Troph | s.e. | Troph | s.e. | ||
| From diet composition | |||||
| From individual food items | |||||
| Ref. | |||||