| Main Ref. | Smith, B.D., 2003 | 
|---|---|
| Remarks | Found in reef areas of lagoons (Ref. 97298); on reef flats and on rocks along the tide line (Ref. 87884). | 
| Marine - Neritic | Marine - Oceanic | Brackishwater | Freshwater | |
|---|---|---|---|---|
| Marine zones / Brackish and freshwater bodies | 
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| Substrate | Benthic: mobile; Soft Bottom: sand; mud; Hard Bottom: rocky; | 
|---|---|
| Substrate Ref. | Springsteen, F.J. and F.M. Leobrera, 1986 | 
| Special habitats | Beds: rock; Coral Reefs: reef flats; lagoons; | 
| Special habitats Ref. | Rehder, H.A., 1980 | 
| Ref. | Smith, B.D., 2003 | 
|---|---|
| Associations | |
| Associated with | |
| Association remarks | |
| Parasitism | 
| Feeding type | mainly animals (troph. 2.8 and up) | 
|---|---|
| Feeding type Ref. | Reichelt, R.E., 1982 | 
| Feeding habit | |
| Feeding habit Ref. | 
| Estimation method | original sample | unfished population | Remark | ||
|---|---|---|---|---|---|
| Troph | s.e. | Troph | s.e. | ||
| From diet composition | |||||
| From individual food items | 3.06 | 0.26 | Tentative trophic level derived from 1 + troph of a single food item | ||
| Ref. | |||||