| Main Ref. | Yeo, D.C.J. and P.K.L. Ng, 1996 | 
|---|---|
| Remarks | Found in well-shaded forest streams above tidal influence with sand and muddy substrates at 0.4 to 1 m depth. Prefers to build burrows along sheltered stream bottoms. | 
| Marine - Neritic | Marine - Oceanic | Brackishwater | Freshwater | |
|---|---|---|---|---|
| Marine zones / Brackish and freshwater bodies | 
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| Substrate | Benthic: mobile; Soft Bottom: sand; mud; | 
|---|---|
| Substrate Ref. | Yeo, D.C.J. and P.K.L. Ng, 1996 | 
| Special habitats | Other habitats: burrows; | 
| Special habitats Ref. | Yeo, D.C.J. and P.K.L. Ng, 1996 | 
| Ref. | |
|---|---|
| Associations | |
| Associated with | |
| Association remarks | |
| Parasitism | 
| Feeding type | |
|---|---|
| Feeding type Ref. | |
| Feeding habit | |
| Feeding habit Ref. | 
| Estimation method | original sample | unfished population | Remark | ||
|---|---|---|---|---|---|
| Troph | s.e. | Troph | s.e. | ||
| From diet composition | |||||
| From individual food items | |||||
| Ref. | |||||