| Main Ref. | Conand, C., 1998 | 
|---|---|
| Remarks | Dwells in moderately shallow waters (despite its English common name), in sandy areas with large amounts of coral rubble; mostly on reef flats of fringing and lagoon-islet reefs; abundant in seagrass beds, on rubble flats, and compact flats where populations can reach high densities (up to 1/m) (Ref. 122); it attaches to rocks with its tube feet (Ref. 118). | 
| Marine - Neritic | Marine - Oceanic | Brackishwater | Freshwater | |
|---|---|---|---|---|
| Marine zones / Brackish and freshwater bodies | 
 | 
 | 
 | 
 | 
| Substrate | Benthic: mobile; Soft Bottom: sand; Hard Bottom: rocky; | 
|---|---|
| Substrate Ref. | |
| Special habitats | Beds: sea grass; Coral Reefs: reef flats; | 
| Special habitats Ref. | Kerr, A.M., K. Netchy and A.M. Gawel, 2006 | 
| Ref. | Conand, C., 1998 | 
|---|---|
| Associations | |
| Associated with | |
| Association remarks | |
| Parasitism | 
| Feeding type | mainly plants/detritus (troph. 2-2.19) | 
|---|---|
| Feeding type Ref. | Conand, C., 1998 | 
| Feeding habit | browsing on substrate | 
| Feeding habit Ref. | Conand, C., 1998 | 
| Estimation method | original sample | unfished population | Remark | ||
|---|---|---|---|---|---|
| Troph | s.e. | Troph | s.e. | ||
| From diet composition | |||||
| From individual food items | |||||
| Ref. | |||||