Bisby, F.A., M.A. Ruggiero, K.L. Wilson, M. Cachuela-Palacio, S.W. Kimani, Y.R. Roskov, A. Soulier-Perkins and J. van Hertum. 2005. (Ref. 19)
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Size / Weight / Age
Climate / Range
Mid-dorsal spines: on segments 6 to 10, uniformly increasing in length posteriorly. Lateral spines: on segments 7 to 12, with adhesive glands at the base of L-4 and L-7. Lateral accessory spines: absent. Lateral terminal spines: long, between 152 to 168 um, 42.2 to 61.7% of trunk length. Abundant prominent cuticular perforation sites arranged in a distinctive pattern. Males possibly lack L-7 and with short seta instead of L-12 spine; lateral terminal accessory spine absent, instead with three pairs of penile spines (Ref. 1409).
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Estimation of some characteristics with mathematical models